1.The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.
研究人员用这些照片来测量老虎脚印的长度和宽度以及脚趾之间的角度，以及10种其他变量来确定不同老虎的“身份”。
2.The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.
研究人员用这些照片来测量老虎脚印的长度和宽度以及脚趾之间的角度，以及10种其他变量来确定不同老虎的“身份”。
3.He says camera traps have identified 24 and the rest have been calculated using the unreliable pugmark method.
据他说，照相机拍摄到24只，而其余的则是依据足迹等不太可靠的方法计算得来。